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Systematics and Biogeography of the "Malvinella Group," Calmoniidae (Trilobita, Devonian)

Bruce S. Lieberman, Gregory D. Edgecombe and Niles Eldredge
Journal of Paleontology
Vol. 65, No. 5 (Sep., 1991), pp. 824-843
Published by: Paleontological Society
Stable URL: http://www.jstor.org/stable/1305809
Page Count: 20
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Systematics and Biogeography of the "Malvinella Group," Calmoniidae (Trilobita, Devonian)
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Abstract

Phylogenetic parsimony analysis forms a basis for classification of the Siegenian--Eifelian "Malvinella group," a monophyletic taxon within the "Metacryphaeus group" of the family Calmoniidae. Fifty-one exoskeletal characters for Metacryphaeus Reed and 12 taxa of the "Malvinella group" yield a shortest length cladogram with a consistency index of 0.51. A classification based on retrieving the structure of this cladogram recognizes six genera: Malvinella Wolfart, Vogesina Wolfart, Palpebrops n. gen. (type P. donegalensis n. sp.), the sister taxa Parabouleia Eldredge and Bouleia Kozlowski, and Plesiomalvinella n. gen. [type P. pujravii (Wolfart)]. Palpebrops is more closely related to Vogesina than to Malvinella; Amazonian species are nested within this group as sister taxa to Andean or Amazon/Andean clades. Bolivian Belén Formation Malvinella buddeae n. sp. is an allopatric species distinct from Icla Formation M. haugi (Kozlowski). "Metacryphaeus" of recent revisions is a paraphyletic grade group including primitive species of the "Malvinella group." Metacryphaeus australis (Clarke), M. tuberculatus (Kozlowski), and M. caffer (Salter) are endemic to the Paraná Basin, Andean Bolivia--Parnaíba Basin, and South Africa, respectively.

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