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Phylogeny of the African Genera Anomatheca and Freesia (Iridaceae: Ixioideae), and a New Genus Xenoscapa

Peter Goldblatt and John C. Manning
Systematic Botany
Vol. 20, No. 2 (Apr. - Jun., 1995), pp. 161-178
DOI: 10.2307/2419447
Stable URL: http://www.jstor.org/stable/2419447
Page Count: 18
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Abstract

Anomatheca and Freesia, African genera of Iridaceae subfamily Ixioideae, are currently thought to be closely related, but until now the exact nature of their alliance has been difficult to determine. Cladistic analysis has provided one means of assessing their relationship. Using 25 characters found to be useful for phylogenetic analysis, including traditional morphological features as well as original data from pollen morphology, leaf anatomy, embryology, and seed development, Freesia appears to be monophyletic but nested in Anomatheca sect. Anomatheca, one of two sections of the genus. A second section, Xenoscapa, which includes one named species and a second one described here, comprises a well-supported clade that is probably only distantly related to Freesia plus sect. Anomatheca. We herein unite Anomatheca and Freesia, but under the later name Freesia. Because Freesia is already a conserved name and because it is a well-known and horticulturally important genus, Anomatheca has been formally rejected in favor of Freesia. We raise sect. Xenoscapa to generic rank and transfer A. fistulosa to it, describe the new species, X. uliginosa, and provide new combinations in Freesia for the species until now included in Anomatheca.

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