If you need an accessible version of this item please contact JSTOR User Support

Angiosperm Flowers and Tricolpate Pollen of Buxaceous Affinity from the Potomac Group (Mid-Cretaceous) of Eastern North America

Andrew N. Drinnan, Peter R. Crane, Else Marie Friis and Kaj Raunsgaard Pedersen
American Journal of Botany
Vol. 78, No. 2 (Feb., 1991), pp. 153-176
Stable URL: http://www.jstor.org/stable/2445239
Page Count: 24
  • Download PDF
  • Cite this Item

You are not currently logged in.

Access your personal account or get JSTOR access through your library or other institution:

login

Log in to your personal account or through your institution.

If you need an accessible version of this item please contact JSTOR User Support
Angiosperm Flowers and Tricolpate Pollen of Buxaceous Affinity from the Potomac Group (Mid-Cretaceous) of Eastern North America
Preview not available

Abstract

A new genus of fossil angiosperms (Spanomera gen. nov.) is established for flowers from two localities in the mid-Cretaceous Potomac Group of Maryland, eastern North America. The type species, Spanomera mauldinensis sp. nov., from the early Cenomanian Elk Neck beds, has inflorescence units with terminal pistillate, and lateral staminate flowers. The organization of inflorescences and flowers is opposite and decussate. Staminate flowers typically have five tepals: two lateral, one posterior, and two in the anterior position. Each tepal is opposed to a stamen with a short filament, dorsifixed anther, and two pairs of pollen sacs. Stamens contain pollen comparable to the dispersed pollen species Striatopollis paraneus (Norris) Singh. Pistillate flowers have two lateral tepals and two anterior-posterior tepals that are opposed to two carpels. Carpels are slightly fused basally along their ventral margins and are semicircular in outline with a long, decurrent, papillate ventral stigma. Frequently this stigmatic surface has abundant attached pollen of the Striatopollis paraneus type. Spanomera marylandensis sp. nov., from the late Albian Patapsco Formation, is similar to S. mauldinensis but is known only from isolated flowers and floral parts. Staminate flowers have four stamens with dorsifixed anthers and each is opposed to a tepal. Stamens contain pollen comparable to the dispersed pollen species Striatopollis vermimurus (Brenner) Srivastava. Carpels have pollen of S vermimurus on the stigma. Spanomera provides further evidence of unisexual but probably insect-pollinated flowers among mid-Cretaceous, early nonmagnoliid ("higher") dicotyledons, and is interpreted as closely related to extant Buxaceae. Characters that Spanomera shares with other taxa suggest that the Buxaceae themselves may be closely related to Myrothamnaceae and other "lower" Hamamelididae.

Page Thumbnails

  • Thumbnail: Page 
153
    153
  • Thumbnail: Page 
154
    154
  • Thumbnail: Page 
155
    155
  • Thumbnail: Page 
156
    156
  • Thumbnail: Page 
157
    157
  • Thumbnail: Page 
158
    158
  • Thumbnail: Page 
159
    159
  • Thumbnail: Page 
160
    160
  • Thumbnail: Page 
161
    161
  • Thumbnail: Page 
162
    162
  • Thumbnail: Page 
163
    163
  • Thumbnail: Page 
164
    164
  • Thumbnail: Page 
165
    165
  • Thumbnail: Page 
166
    166
  • Thumbnail: Page 
167
    167
  • Thumbnail: Page 
168
    168
  • Thumbnail: Page 
169
    169
  • Thumbnail: Page 
170
    170
  • Thumbnail: Page 
171
    171
  • Thumbnail: Page 
172
    172
  • Thumbnail: Page 
173
    173
  • Thumbnail: Page 
174
    174
  • Thumbnail: Page 
175
    175
  • Thumbnail: Page 
176
    176