Behaviour publishes original research pursuing Tinbergen's four questions and questions resulting from the interrelationship among the four. In addition, the editorial board encourages reviews of behavioural biology that illuminate emergent trends and new directions in behavioural research. Niko Tinbergen (1907-1988) defined 4 questions for research in behavioral biology: Proximate causation of behaviour: 1. How does an animal use its sensory and motor abilities to activate and modify its behaviour patterns? (physiological mechanisms) 2. How does an animal's behaviour change during its growth, especially in response to the experiences that it has while maturing? (ontogeny of development) Ultimate causation of behaviour: 3. How does the behaviour promote an animal's ability to survive and reproduce? (adaptation) 4. How does an animal's behaviour compare with that of other closely related species, and what does this tell us about the origins of its behavior and the changes that have occurred during the history of the species? (phylogeny) Niko Tinbergen shared, with Konrad Lorenz and Karl von Frisch, the 1973 Nobel Prize for Medicine or Physiology for contributions to the study of behavioural biology. Tinbergen was at heart an experimentalist who, more than Lorenz and von Frisch, applied the scientific method to the field of animal and human behaviour. It is his experimental approach to the study of behaviour that lasts to this day. That is why Tinbergen listed questions and not answers (theorems or laws). The answers (or at least some of them) are published monthly in Behaviour, the journal Tinbergen co-founded with W. H. Thorpe in 1948.

BRILL, founded in 1683, is a publishing house with a strong international focus. BRILL is renowned for its publications in the following subject areas; Asian Studies, Ancient Near East & Egypt, Biblical Studies & Religious Studies, Classical Studies, Medieval & Early Modern Studies, Middle East & Islamic Studies. BRILL's mainly English language publications include book series, individual monographs and encyclopaedias as well as journals. Publications are increasingly becoming available in electronic format (CD-ROM and/or online editions).BRILL is proud to work with a broad range of scholars and authors and to serve its many customers throughout the world. Throughout its existence the company has been honored with many awards which recognise BRILL's contribution to science, publishing and international trade.